Coates, B.J. (1985) The birds of Papua New Guinea Vol. 1. Non-Passerines. Dove Publications, Australia.
Colonies Surveyed (9)
Papua New Guinean EEZ -
Colony: New Ireland
Papua New Guinean EEZ -
Colony: Kavieng
Papua New Guinean EEZ -
Colony: Bavo Island
Papua New Guinean EEZ -
Colony: Witu Island
Papua New Guinean EEZ -
Colony: Nuguria
Papua New Guinean EEZ -
Colony: Manus
Papua New Guinean EEZ -
Colony: Emirau
Papua New Guinean EEZ -
Colony: Duke of York
Papua New Guinean EEZ -
Colony: Duke of York